HEIDELBERG, Miss. — Three monkeys remained on the loose Wednesday as searchers in masks, face shields and other protective gear scoured fields along a rural Mississippi highway where a truck carrying the primates overturned a day earlier.

It remained unclear who owns the monkeys, who was transporting them and where they were being taken when the truck crashed on Interstate 59 north of Heidelberg. Authorities have said most of the 21 monkeys were killed but haven’t elaborated on just what occurred.

Officials initially said all but one of the escaped monkeys had been killed, leaving one on the loose. But the Jasper County Sheriff’s Department later said officials from Tulane University examined the trailer Tuesday evening and determined three monkeys had escaped.

The truck was carrying

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