(Bloomberg/Sidhartha Shukla and Suvashree Ghosh) — Cryptocurrency traders saw more than $1.5 billion in bullish wagers liquidated on Monday, triggering a sharp selloff that hit smaller tokens hardest.

Ether slumped as much as 9% to $4,075 as nearly half a billion dollars of leveraged long positions in the second-largest token were liquidated, according to data from Coinglass. Bitcoin declined 3% to $111,998 at one point. Coins like Solana, Algorand and Avalanche also slipped.

Demand from publicly-listed vehicles set up to hoard tokens helped drive Bitcoin and Ether to all-time highs in August. That momentum has started to fade as shares of digital-asset treasury firms ranging from Michael Saylor’s Strategy to Japan’s Metaplanet Inc. retreated.

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