Jane Goodall, the British conservationist and primatologist renowned for her groundbreaking research on chimpanzees, has died aged 91.

The Jane Goodall Institute announced in a Facebook post on Wednesday that Goodall died of natural causes in California during a speaking tour in the United States.

“Dr Goodall’s discoveries as an ethologist transformed science, and she was a tireless advocate for the protection and restoration of the natural world,” the institute said.

Born in London in 1934, Goodall began researching free-living chimpanzees in Tanzania in 1960.

She observed a chimpanzee named David Greybeard make a tool from twigs and use it to fish termites from a nest, a ground-breaking observation that challenged the definition of humans as the single species capable of making tools

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