James D. Watson, the controversial American biologist whose 1953 discovery of the structure of DNA ushered in the age of genetics, has died at the age of 97.

His work provided the foundation for the biotechnology revolution of the late 20th century.

His death was confirmed by Cold Spring Harbor Laboratory on Long Island, where he worked for many years.

The New York Times reported Mr Watson died this week at a hospice on Long Island.

In his later years, Mr Watson's reputation was tarnished by comments on genetics and race that led him to be ostracised by the scientific establishment.

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