SAN DIEGO, California — Gramma, a Galápagos tortoise and the oldest known resident of the San Diego Zoo, has died at an estimated age of 141, CBS News reported .

Zoo officials said Gramma died on Nov. 20 after caretakers determined her declining bone condition, attributed to advanced age, made continued care unsustainable, according to the Los Angeles Times .

Born in her native Galápagos habitat and later arriving at the zoo between 1928 and 1931 as part of the facility’s first Galápagos tortoise group, she became a quiet but major fixture of the zoo grounds, 10 News San Diego reported .

In her long lifetime, the TV station said Gramma lived through more than two World Wars, more than 20 U.S. presidencies and vast global shifts, all while delighting visitors, young and old, who wa

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